Sometimes it's well worth the extra $$ to go to the better model, sometimes not so much. As you say, sometimes you're paying for a little more smoothness in cranking and not much else, other times you're getting significant functional differences.
I did a back-to-back review between the 8000OC and 8000D on my Facebook page a while ago -
https://www.facebook.com/reelguts/photos/?tab=album&album_id=1554919904815379
I started that page to give fishos a look into what's actually going on inside reels and what their dollars are buying them, is it just a marketing term and a brand or are you actually getting an improved product?
Conclusion was that the 8000D is absolutely worth the extra $50, but the final conclusion was that the Thunnus is the one to go for. At $269 that's a great price for the Thunnus, the Ci4 rotor is key as the D/OC rotor (identical) flexes too much at more than ~6kg drag in the 8000 size. Off the top of my head the Thunnus gets the hardened spool lip as well, which may also be smoother = longer casts? Quite a few subtle little changes in the Thunnus, important ones are the spool bearing, line roller bearing, "Paladin" gears and the obvious aforementioned material change.
In some other reels... yes and no. Read Alan Hawk on Shimano Saragosa SW vs Spheros SW as a good example.
Daiwa BG is a fantastic "budget" reel, as is the Okuma Azores.